躍る

What does 躍る mean in English?

1. to jump up; to spring up; to leap(v5r, vi)
2. to move around; to bounce up and down(v5r, vi)
3. to pound (of one's heart, e.g. with excitement); to throb(v5r, vi)
4. to be messy (of handwriting); to be untidy(v5r, vi)